Life and health

Harold E. Varmus

Harold E. Varmus (born 1939) is an American oncologist who shared the 1989 Nobel Prize with J. Michael Bishop for discovering the cellular origin of retroviral oncogenes, and later directed NIH, Memorial Sloan Kettering, and the NCI.

Harold Edwin Umbarger

Harold Edwin Umbarger (1921–1999) was an American biochemist and geneticist at Purdue University who discovered feedback inhibition of enzymes and defined how bacteria synthesize the branched-chain amino acids.

Harold Erickson

Harold Erickson is a cell biologist and James B. Duke Distinguished Professor Emeritus at Duke University, known for his 1995 proposal that bacterial FtsZ is a tubulin homolog.

Harold F. Dvorak

Harold F. Dvorak is an American pathologist and vascular biologist at Harvard Medical School who discovered VEGF, the key growth factor behind tumor angiogenesis, in 1983.

Harold Gainer

Harold (Hal) Gainer (died 2023) was an American neuroscientist who worked at NIH from 1969 to 2012, known for work on neurosecretion and posterior pituitary hormones.

Harold Ginsberg

Harold Samuel Ginsberg was an American virologist who showed that adenoviruses cause acute respiratory disease and childhood pneumonia, and chaired microbiology at Penn and Columbia.

Harold I. Feldman

Harold I. Feldman is an American physician-epidemiologist at the University of Pennsylvania who studies kidney disease, led national NIH-funded kidney research networks, and edits the American Journal of Kidney Diseases.

Harold J. Burstein

Harold J. Burstein is an American medical oncologist specializing in breast cancer, a professor at Harvard Medical School, and a Dana-Farber clinician known for NEJM reviews and treatment guidelines.

Harold J. Evans

Harold J. Evans (1921–2007) was an American plant physiologist at Oregon State University who pioneered the study of mineral nutrition in plants and biological nitrogen fixation.

Harold J. Fallon

Harold J. Fallon (1931–2018) was an American hepatologist who chaired medicine at the Medical College of Virginia and led the landmark corticosteroid trial for alcoholic hepatitis.

Harold J. Wanebo

Harold J. Wanebo was an American surgical oncologist at Roger Williams Medical Center in Rhode Island, known for linking blood tests and tumor depth to cancer prognosis.

Harold Kelley

Harold Kelley (1921–2003) was an American social psychologist at UCLA, best known for his covariation model of causal attribution and the interdependence theory he developed with John Thibaut.

Harold L. Moses

Harold L. Moses is an American cancer biologist who discovered that TGF-β inhibits cell proliferation and founded and directed the Vanderbilt-Ingram Cancer Center from 1993 to 2005.

Oct 10, 2026

Harold Leeming Sheehan

Harold Leeming Sheehan (1900–1988) was a British pathologist who, working from autopsies in Glasgow, showed in 1937 that postpartum hemorrhage and shock can destroy the anterior pituitary, the condition now called Sheehan's syndrome.

Oct 10, 2026

Harold Munro Fox

Harold Munro Fox, born Harold Munro Fuchs, was a British zoologist known for work on invertebrate respiratory pigments and for his forty-year editorship of Biological Reviews.

Harold O. Conn

Harold O. Conn (1925–2011) was an American hepatologist at Yale and the West Haven VA hospital who coined "spontaneous bacterial peritonitis" and created the West Haven criteria for hepatic encephalopathy.

Harold P. Freeman

Harold P. Freeman was an American surgical oncologist who pioneered patient navigation for poor and uninsured cancer patients, founding the first navigator program in Harlem in 1990.

Harold R. Collard

Harold R. Collard is an American pulmonologist and physician-scientist who has been Vice Chancellor for Research at the University of California, San Francisco since 2022, known for interstitial lung disease and pulmonary fibrosis research.

Oct 10, 2026

Harold Raistrick

Harold Raistrick (1890–1971) was a British biochemist, internationally acknowledged pioneer of fungal product chemistry, who held the biochemistry chair at the London School of Hygiene and Tropical Medicine from 1929.

Harold S. Margolis

Harold S. Margolis was an American CDC medical epidemiologist who led federal viral hepatitis research and was senior author of the landmark 1999 national hepatitis C prevalence study.

Harold T. Shapiro

Harold T. Shapiro, born 1935, is a Canadian-born American economist who was president of the University of Michigan from 1980 to 1988 and Princeton's 18th president from 1988 to 2001.

Harold W. Jaffe

Harold W. Jaffe is an American physician and epidemiologist who joined the CDC's 1981 AIDS task force, led its first national case-control study, and directed its HIV center.

Harold Weintraub

Harold Weintraub (1945–1995) was an American molecular biologist at the Fred Hutchinson Cancer Research Center in Seattle who identified MyoD, the single gene that triggers muscle differentiation in non-muscle cells.

HARP (algorithm)

HARP (harmonic phase) MRI algorithm, by Osman and Prince, Johns Hopkins, 1999: rapid analysis of SPAMM-tagged cardiac MR images using invariant harmonic phase to track myocardial motion and strain; basis of Diagnosoft HARP software.

Harp seal

Harp seal (Pagophilus groenlandicus), also called Greenland seal or saddle-back seal: an ice-breeding phocid seal of the North Atlantic with a harp-shaped back patch, whitecoat pups, three populations, and a long commercial hunt history.

Harpacticoida

Harpacticoida, an order of benthic copepod crustaceans (G. O. Sars, 1903; harpacticoid copepods), with 463 genera and about 3,000 species in marine, freshwater, and moist terrestrial habitats; key meiofauna, fish prey, and live feed.